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The world has entered a biodiversity crisis. The rapid loss of species as a consequence of human alteration of the biosphere has attracted widespread attention among ecologists and spawned a voluminous literature concerning the cataloguing, description, prediction, rescue, and management of biodiversity. Marine ecologists have long been interested in explaining patterns in community diversity, especially in the benthos. Much of this effort has been directed at deciphering the role of biological interactions, such as competition, in maintaining diverse communities, and this debate has perhaps been most pronounced amongst workers interested in deep sea-floor and rocky shore environments. However, it has been suggested that competition is relatively unimportant in intertidal....